Click here for a PDF list of publications, here for my publications on Google Scholar, here for my CV, or on the download-icon below to download individual papers.

Books (as author)

Ramsey, G. (2023) Human Nature. Cambridge University Press. Free to download here.
Ramsey, G. and Durand, P. (expected 2025) Death and Taxa. Cambridge University Press.
Fábregas-Tejeda, A. and Ramsey, G. (expected 2025) Niche Construction and Evolutionary Theory. Cambridge University Press.
Durand, P. and Ramsey, G. (expected 2026) Evolution’s End: Life, Death, and the Mortality Constraint. Oxford University Press.

Books (as editor)

Ramsey, G. and Aaby, B. (expected 2026) Psychodiversity: Cognition and Sentience Beyond Humans. Routledge.
Desmond, H. and Ramsey, G. (2023) Human Success: Evolutionary Origins and Ethical Implications. Oxford University Press. For more information, visit the Oxford University Press website or purchase it via Amazon
Ramsey, G. and De Block, A. (2022) The Dynamics of Science: Computational Frontiers in History and Philosophy of Science. University of Pittsburgh Press. For more information, visit the University of Pittsburgh Press website or purchase it via Amazon
Ramsey, G. and Pence, C. H. (2016) Chance in Evolution. University of Chicago Press. For more information, visit the University of Chicago Press website or purchase it via Amazon

Articles and book chapters

download-icon Eronen, M. I. and Ramsey, G. (forthcoming) “What are the ‘levels’ in levels of selection?” The British Journal for the Philosophy of Science. DOI: 10.1086/718990
Durand, P. and Ramsey, G. (forthcoming) “Conceptual Foundations of Cell Mortality” in J. M. Hardwick and P. Durand (eds.) The New Era of Microbial Cell Death Springer.
download-icon Fábregas-Tejeda, A. and Ramsey, G. (2024) “Driftability and niche construction” Synthese 204: 162. DOI: 10.1007/s11229-024-04815-5
download-icon Ramsey, G. and Villegas, C. (2024) “Developmental Channeling and Evolutionary Dappling” Philosophy of Science 91(4): 869-886. DOI: 10.1017/psa.2024.5
download-icon Aaby, B., Dani, G. and Ramsey, G. (2024) “Explanatory Gaps in Evolutionary Theory” Biology & Philosophy 39: 22. DOI: 10.1007/s10539-024-09957-x
download-icon Meneganzin, A., Ramsey, G, and DiFrisco, J. (2024) “What is a Trait? Lessons from the Human Chin” Journal of Experimental Zoology Part B: Molecular and Developmental Evolution 343(2): 65-75. DOI: 10.1002/jez.b.23249
download-icon DiFrisco, J. and Ramsey, G. (2023) “Adaptationism and trait individuation” Philosophy of Science 90(5): 1234-1243. DOI: 10.1017/psa.2023.28
download-iconRamsey, G. and Durand, P. (2023) “Cell Fate: What’s Evolution Got to do With It?” Yale Journal of Biology and Medicine 96(4): 565-568. DOI: 10.59249/FBHI3484
Desmond, H. and Ramsey, G. (2023) “The Manifold Challenges to Understanding Human Success” in H. Desmond and G. Ramsey (eds.) Human Success: Evolutionary Origins and Ethical Implications, Oxford University Press.
download-icon Durand, P. and Ramsey, G. (2023) “The concepts and origins of cell mortality” History and Philosophy of the Life Sciences 45:23. DOI: 10.1007/s40656-023-00581-8
download-icon Ramsey, G. and Aaby, B. (2022) “The proximate-ultimate distinction and the active role of the organism in evolution” Biology & Philosophy 37: 31. DOI: 10.1007/s10539-022-09863-0
download-icon Ramsey, G. and Deem, M. (2022) “Empathy and the evolutionary emergence of guilt” Philosophy of Science 89(3): 434-453. DOI: 10.1017/psa.2021.36
download-icon Aaby, B. and Ramsey, G. (2022) “Three kinds of niche construction” The British Journal for the Philosophy of Science 73(2): 351-372. DOI: 10.1093/bjps/axz054
De Block, A. and Ramsey, G. (2022) “Tools, tests, and data: An overview of the new history and philosophy of science” in G. Ramsey and A. De Block (eds.) The Dynamics of Science: Computational Frontiers in History and Philosophy of Science, University of Pittsburgh Press.
download-icon Ndhlovu, A., Durand, P., and Ramsey, G. (2021) “Programmed cell death as a black queen in microbial communities” Molecular Ecology 30: 1110-1119. DOI: 10.1111/mec.15757
download-icon Climenhaga, N., DesAutels, L., and Ramsey, G. (2021) “Causal inference from noise” Noûs 55: 152–170. DOI: 10.1111/nous.12300
download-icon Durand, P. and Ramsey, G. (2019) “The nature of programmed cell death” Biological Theory 14: 30–41. DOI: 10.1007/s13752-018-0311-0
download-icon Pence, C. and Ramsey, G. (2018) “How to do digital philosophy of science” Philosophy of Science 58: 930-941. DOI: 10.1086/699697
download-icon Ramsey, G. (2018) “Trait bin and triat cluster accounts of human nature” In: E. Hannon and T. Lewens (eds.) Why We Disagree about Human Nature. Oxford University Press. DOI:10.1093/oso/9780198823650.003.0003
download-icon Ramsey, G. and De Block, A. (2017) “Is cultural fitness hopelessly confused?” The British Journal for the Philosophy of Science 68: 305-328. DOI: 10.1093/bjps/axv047
download-iconRamsey, G. (2017) “What is human nature for?” In: A. Fuentes and A. Visala (ed.) Verbs, Bones and Brains: Interdisciplinary Perspectives on Human Nature, Notre Dame: University of Notre Dame Press, 217-230.
download-icon Ramsey, G. (2017) “What is animal culture?” In: K. Andrews and J. Beck (eds.) Routledge Companion to the Philosophy of Animal Minds. Routledge Press, 345-353.
download-icon Ramsey, G. (2016) “The causal structure of evolutionary theory” Australasian Journal of Philosophy 94: 421-434. DOI: 10.1080/00048402.2015.1111398
download-icon Ramsey, G. and Pence, C. H. (2016) “evoText: A new tool for analyzing the biological sciences.” Studies in History and Philosophy of Biological and Biomedical Sciences 57: 83-87. DOI: 10.1016/j.shpsc.2016.04.003
Ramsey, G. and Pence, C. H. (2016) “Chance in evolution from Darwin to contemporary biology.” In: G. Ramsey and C.H. Pence (eds.) Chance in Evolution. Chicago: University of Chicago Press, 1-11.
download-icon Deem, M. and Ramsey, G. (2016) “Guilt by association?” Philosophical Psychology 29(4): 570-585. DOI: 10.1080/09515089.2015.1126706
download-icon Ramsey, G. (2016) “Can altruism be unified?” Studies in History and Philosophy of Biological and Biomedical Sciences 56: 32-38. DOI: 10.1016/j.shpsc.2015.10.007
download-icon De Block, A. and Ramsey, G. (2016) “The organism-centered approach to cultural evolution.” Topoi 35: 283-290. DOI: 10.1007/s11245-014-9283-2
download-icon Deem, M. and G. Ramsey (2016) “The Evolutionary Puzzle of Guilt: Individual or Group Selection?” Emotion Researcher, ISRE’s Sourcebook for Research on Emotion and Affect, Andrea Scarantino (ed.), http://emotionresearcher.com/the-evolutionary-puzzle-of-guilt-individual-or-group-selection/
download-icon Pence, C. and Ramsey, G. (2015) “Is organismic fitness at the basis of evolutionary theory?” Philosophy of Science 82: 1081–1091. DOI: 10.1086/683442
download-iconRamsey, G. (2013) “Human nature in a post-essentialist world.” Philosophy of Science 80: 983-993. DOI: 10.1086/673902.
download-icon Ramsey, G. (2013) “Can fitness differences be a cause of evolution?” Philosophy & Theory in Biology 5:e401. http://hdl.handle.net/2027/spo.6959004.0005.001.
download-icon Pence, C.H. and Ramsey, G. (2013) “A new foundation for the propensity interpretation of fitness.” The British Journal for the Philosophy of Science 64:851-881. DOI: 10.1093/bjps/axs037.
download-icon Ramsey, G. (2013) “Organisms, traits, and population subdivisions: two arguments against the causal conception of fitness?” The British Journal for the Philosophy of Science 64: 589-608 DOI: 10.1093/bjps/axs010.
download-icon Ramsey, G. (2013) “Culture in humans and other animals.” Biology and Philosophy 27: 457-479. DOI: 10.1007/s10539-012-9347-x.
download-icon Ramsey, G. (2013) “Driftability.” Synthese 190: 3909-3928. DOI: 10.1007/s11229-012-0232-6.
download-icon Ramsey, G. (2012) “How human nature can inform human enhancement.” Philosophy and Technology 25: 479-483. DOI: 10.1007/s13347-012-0087-2.
download-icon Ramsey, G. and Peterson, A. (2012) “Sameness in biology.” Philosophy of Science 77: 255-275. DOI: 10.1086/664744.
download-icon Ramsey, G. and Brandon, R. (2011) “Why reciprocal altruism is not a kind of group selection.” Biology and Philosophy 26: 385-400. DOI: 10.1007/s10539-011-9261-7.
download-icon Ramsey, G. (2007) “The Fundamental Constraint on the evolution of culture.” Biology and Philosophy 22: 401-414. DOI: 10.1007/s10539-006-9038-6.
download-icon Ramsey, G., Bastian, M. L., and van Schaik, C. (2007) “Animal innovation defined and operationalized.” Behavioral and Brain Sciences 30: 393-437. DOI: 10.1017/S0140525X07002373.
download-icon Brandon, R. and Ramsey, G. (2007) “What’s Wrong with the Emergentist Statistical Interpretation of Natural Selection and Random Drift?” In: D. Hull and M. Ruse (eds.) Cambridge Companion to the Philosophy of Biology, Cambridge: Cambridge University Press.
download-icon Ramsey, G. (2006) “Block fitness.” Studies in History and Philosophy of Biological and Biomedical Sciences 37: 484-498. DOI: 10.1016/j.shpsc.2006.06.009.
download-icon Cobb, A., Nadkarni, N., Ramsey, G., and Svoboda, A. J. (2001) “Recolonization of bigleaf maple branches by epiphytic bryophytes following experimental disturbance.” The Canadian Journal of Botany 79: 1-8. DOI: 10.1139/b00-134

Encyclopedia Entries and Bibliographies

download-icon Ramsey, G. and Desmond, H. (2017) “Philosophy of Biology” Oxford Bibliography.
download-icon Ramsey, G. and Pence, C. H. (2013) “Fitness: philosophical problems.” In: eLS. Chichester: John Wiley & Sons, Ltd. DOI: 10.1002/9780470015902.a0003443.pub2

Book reviews

download-icon Hollocher, H., Pence, C. H., Ramsey, G., and Wirth, M. M. (2013) “A path to success? A Review of Evolution, Development, and the Predictable Genome by David L. Stern.” Evolution and Development 15(1): 80–82. DOI: 10.1111/ede.12016.
download-icon Hollocher, H., Fuentes, A., Pence, C. H., Ramsey, G., Sportiello, D. J., and Wirth, M. M. (2011) “[Review of] On the Origin of Stories: Evolution, Cognition, and Fiction.” The Quarterly Review of Biology 86(2): 137-138. DOI: 10.1086/659913.
download-icon Pence, C. H., Hollocher, H., Nichols, R., Ramsey, G., Siu, E., and Sportiello, D. J. (2011) “[Review of] Did Darwin Write the Origin Backwards? Philosophical Essays on Darwin’s Theory.” Philosophy of Science 78(4): 705-709. DOI: 10.1086/661775.
download-icon Ramsey, G., Hollocher, H., Fuentes, A., Pence, C. H., and Siu, E. (2010) “[Review of] Darwinian Populations and Natural Selection.” The Quarterly Review of Biology 85(4): 499-500. DOI: 10.1086/656856.

Blog and magazine articles

Ramsey, G. (2023, May 04) “The culture of bees and the AI apocalypse” IAI News.
De Block, A. and Ramsey, G. (2015) “The life of Culture” OUP Blog.

Books (as illustrator)

Lombardi, A. (1998) Respecting the Knowledge: Ethnobotany of Western Washington. Olympia: Washington State Capital Museum. (Illustrated by Grant Ramsey.)

Other miscellaneous publications

Ramsey, G., Longino, J., and Kugler, C. (1998) Formicidae: Rogeria creightoni. https://ants.biology.utah.edu/GENERA/ROGERIA/SPECIES/creightoni/creightoni.html. (One of 13 published web pages on the ant genus Rogeria. See full list of species here: https://ants.biology.utah.edu/GENERA/ROGERIA/Specieslist.html)